When the complex in photosystem II is activated by light energy, it gives up electrons, which are then absorbed by an electron acceptor. This electron acceptor is part of what is called an energy transfer system, through which the electrons move until they reach photosystem I. During these transfers, hydrogen ions are pumped from the stroma into the interior of the thylakoid. As hydrogen ions lead back across the membrane through special carrier proteins called ATPases, ADP is phosphorylated, forming ATP (ADP → ATP).